The Wheeler Shale (named by Charles Walcott) is a Cambrian (c.507 Ma) fossil locality world famous[1] for prolific agnostid and Elrathia kingii trilobite remains (even though many areas are barren of fossils)[2] and represents a Konzentrat-Lagerstätten. Varied soft bodied organisms are locally preserved, a fauna (e.g. Naraoia, Wiwaxia & Hallucigenia) and preservation style (carbonaceous film) normally associated with the more famous Burgess Shale.[3] As such, the Wheeler Shale also represents a Konservat-Lagerstätten.[4] Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms 490 to 610m (1,600 to 2,000ft) of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America.[5] At the type locality of Wheeler Amphitheater, House Range, Millard County, west Utah the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thin, flaggy limestone.[6] The Wheeler Formation (although the Marjum & Weeks Formations are missing) extends into the Drum Range, N.W. of the House Range where similar fossils and preservation are found.[6] Detailed work recognises a number of ~10 m thick lagerstätten sequences in the formation, each of which formed at a sea-level high stand[7] in deep water.[8] The lagerstätte were deposited by turbitides and mudslides onto an oxygenated sea floor.[7] Recorded organisms include brachiopods, primitive echinoderms, sponges, chancellorids, carpoids, eocrinoids (e.g. Gogia), trilobitomorphs, varied arthropods (e.g. Anomalocaridids, Emeraldella, Phyllocarids, trilobites), annelid & priapulid worms, jelly fish & algae.[9][6][10][11][12][13][14] Intriguingly, the preservation of hard bodied trilobite remains and soft bodied animals seems to be mutually exclusive within particular horizons.[15][4] [edit] Gallery Asaphiscus wheeleri, Cambrian, Wheeler shale, Utah. | | [edit] References - ^ Johnson, Kirk; Troll, Ray (2007), Cruising the fossil freeway: An epoch tale of a scientist and an artist on the ultimate 5,000-Mile paleo road trip, Golden, CO.: Fulcrum Publishing, ISBN 978-1-55591-451-6
- ^ Robert R. Gaines; Mary L. Droser (2003). "Paleoecology of the familiar trilobite Elrathia kingii: An early exaerobic zone inhabitant" (pdf). Geology 31: 941-4. http://earthsciences.ucr.edu/docs/Gaines&Droser_2003.pdf.
- ^ Robert R. Gaines; Derek E.G. Briggs ; Zhao Yuanlong (2008). "Cambrian Burgess Shale–type deposits share a common mode of fossilization". Geology 36: 755-758. doi:10.1130/G24961A.1.
- ^ a b Gaines, R (2005). "A New Hypothesis for Organic Preservation of Burgess Shale Taxa in the Middle Cambrian Wheeler Formation, House Range, Utah". Palaeogeography Palaeoclimatology Palaeoecology 220: 193-205. doi:10.1016/j.palaeo.2004.07.034.
- ^ Robison, R.A. (1964), "Late Middle Cambrian faunas from western Utah", Journal of Paleontology 38: 510-566, http://jpaleontol.geoscienceworld.org/cgi/content/abstract/38/3/510
- ^ a b c Gunther, L.F.; Gunther, V.G. (1981), "Some Middle Cambrian Fossils of Utah", Brigham Young University Geology Studies 28: 1-81
- ^ a b Brett, C. E.; Allison, P. A.; Desantis, M. K.; Liddell, W. D.; Kramer, A. (2009). "Sequence stratigraphy, cyclic facies, and lagerstätten in the Middle Cambrian Wheeler and Marjum Formations, Great Basin, Utah". Palaeogeography Palaeoclimatology Palaeoecology 277: 9–33. doi:10.1016/j.palaeo.2009.02.010. edit
- ^ Halgedahl, S. L.; Jarrard, R. D.; Brett, C. E.; Allison, P. A. (2009). "Geophysical and geological signatures of relative sea level change in the upper Wheeler Formation, Drum Mountains, West-Central Utah: A perspective into exceptional preservation of fossils". Palaeogeography Palaeoclimatology Palaeoecology 277 (1-2): 34–56. doi:10.1016/j.palaeo.2009.02.011. edit
- ^ Briggs, D.E.G.; Robison, R.A. (1984), Exceptionally preserved nontrilobite arthropods and Anomalocaris from the Middle Cambrian of Utah, The Paleontological Institute, The University of Kansas, http://kuscholarworks.ku.edu/dspace/handle/1808/3656
- ^ Photos of Wheeler Shale fossils from UC Berkeley
- ^ Rigby, J.K. (1978), "Porifera of the Middle Cambrian Wheeler Shale, from the Wheeler Amphitheater, House Range, in western Utah.", Journal of Paleontology 52: 1325-1345
- ^ Utah's Cambrian Life from University of Kansas Natural History Museum
- ^ Cambrian fossils from Utah by the University of Utah
- ^ Comprehensive treatment from The Virtual Fossil Museum
- ^ Gaines, Robert R.; Droser, Mary L.; Kennedy, Martin J. (2001), "Taphonomy of soft-bodied preservation and ptychopariid Lagerstätte in the Wheeler Shale (Middle Cambrian), House Range, USA; controls and implications.", PaleoBios 21 (Suppl.2): 1-55, http://www.ucmp.berkeley.edu/napc/abs9.html
| Modes of preservation in the Cambrian | | | Exceptional | | Phosphate | | | | | | | | Carbonaceous film | | | | | | | | Casts and moulds | | |  Legged trilobite fossil | | | Conventional | | | |