A muscle spindle, with γ motor and Ia sensory fibers

Type Ia Sensory Fiber also called Primary Afferent Fiber is a type of sensory fiber. It is a component of a muscle fiber's muscle spindle which keeps track of how fast a muscle stretch changes (the velocity of the stretch).

[edit] Function of muscle spindles

In order to control movements, the nervous system must receive continuous sensory information from muscles and joints. For this purpose the body has specialized sensory receptors called proprioceptors. Muscle spindles are a type of proprioceptor, and they are located inside the muscle itself. They are sensitive to muscle length because they are in parallel with the contractile fibers.

[edit] Types of sensory fibers

This change in length of the spindle is transduced (transformed into electric membrane potentials) by two types of sensory afferents, whose cell bodies are located in dorsal root ganglia located next to the spinal cord.

The two kinds of sensory fibers are different in respect to the kind of potentials they generate:

The first of the two main groups of stretch receptors wrapping the intrafusal fibers are the Ia fiber, which are the largest and fastest fibers, and they fire when the muscle is stretching. They are characterized by their rapid adaptation, because as soon as the muscle stops changing length, the Ia stop firing and adapt to the new length. Ia fibers essentially supply proprioceptive information about the rate of change of its respective muscle: the derivative of the muscle's length (or position).

Type 1a fibers connect to both nuclear bag fibers and nuclear chain fibers. These connections are also called "annulospiral endings ".

[edit] Efferent innervation

In addition, the spindle also has a motor efferent innervation carried by gamma motor neurons, which is used by the nervous system to modify the spindle's sensitivity.

[edit] Termination of afferents

Ia afferents from the muscle spindle terminate on the proximal dendrites of motor neurones.

[edit] See also

• Intrafusal muscle fiber
• Proprioception
• Motor system
• Muscle
• Muscle spindle
• Reflex action
• Phasic stretch reflex
• Tonic stretch reflex
• Type II sensory fiber

[edit] External links

• Lecture notes from John D.C. Lambert on neurophysiology.
• http://www.lib.mcg.edu/edu/eshuphysio/program/section8/8ch3/s8ch3_15.htm