Proton gradient: Solid green: sieve tube; dashed green: sieve tube plates; light pink: companion cell; dark pink: nucleus; yellow: nutrients

In plant anatomy, sieve vascular tissue tube elements, also called sieve tube members, are a type of elongated parenchyma cells in phloem tissue. At the ends these cells are connected with other sieve elements, and together they constitute the sieve tube. The main function of the sieve tube is transport of carbohydrates in the plant (e.g., from the leaves to the fruits and roots). Unlike vessel elements, which are elongated cells that transport water and minerals in the xylem/wood that are dead when mature, and represent another kind of vascular tissue in the plant, sieve elements are living cells. They are thick and circular and can be different colours. Sieve tubes are not known as 'Real Cells' as they lack a substantial amount of Cytoplasm.

At the interface between two sieve tube members in angiosperms are sieve plates, pores in the plant cell walls that facilitate the movement of liquid. Neighbouring each of the sieve tube elements is a minimum of one companion cell, connected by plasmodesmata (channels between the cells). Sieve tube members have no cell nucleus, ribosomes, or a vacuole, the nucleus and ribosomes of its companion cell(s) compensate for this. In leaves, these cells help in moving the sugar produced by photosynthesis in the mesophyll tissue into the sieve tube elements. Sieve tubes are mainly to transport sugars and nutrients up and down the plant.

Sieve cells are long, slender, conducting cells of the secondary phloem that do not form a constituent element of a sieve tube, but which are provided with relatively unspecialized sieve areas, especially in the tapering ends of the cells that overlap those of other sieve cells. They have a narrower diameter and are more elongated compared to sieve tube members, the other kind of sieve elements present in the phloem. Sieve cells are associated with albuminous cells, which lack starch, thus making it possible to differentiate them from phloem parenchyma.

The forest botanist Theodor Hartig was the first to discover and name these cells as Siebfasern (sieve fibres) and Siebröhren (sieve tubes) in 1837.

[edit] See also

• Phloem
• Vascular tissue

[edit] External links

• Comprehensive explanation of sieve elements - University of Hamburg Department of Biology
• Introduction to secondary phloem tissue by Dr. David T. Webb - University of Hawaii

[edit] References

• Katherine Esau (1969). The Phloem -in: Encyclopedia of Plant Anatomy. Gebrüder Borntraeger, Berlin, Stuttgart.
• Campbell, N. A. (1996) Biology (4th edition). Benjamin Cummings NY. ISBN 0-8053-1957-3
• Hartig, T. (1837) Vergleichende Untersuchungen über die Organisation des Stammes der einheimischen Waldbäume. Jahresber. Fortschr. Forstwiss. Forstl. Naturkd. 1:125-168