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Eleutherozoa
Pisaster ochraceaus
Scientific classification
Kingdom: Animalia
Phylum: Echinodermata
Subphylum: Eleutherozoa
Bather, 1900
Superclasses

Eleutherozoa is a subphylum of echinoderms. They are mobile animals with the mouth directed towards the substrate. They usually have a madreporite, tube feet, and moveable spines of some sort, and some have Tiedemann's bodies on the ring canal. All living echinoderms except Crinoidea belong here.

[edit] Systematics

There are 2 main competing hypotheses about the internal subdivision, both about equally well supported by both molecular and morphological data. They differ in their placement of the Ophiuroidea (brittle stars), and are named accordingly.

The "Cryptosyringida" hypothesis posits that the "sea-star" morphology is plesiomorphic for Eleutherozoa as a whole, and that starfish (Asteroidea) and brittle stars are not very closely related, the latter forming the clade Cryptosyringida together with the Echinozoa. The "Asterozoa" hypothesis on the other hand implies that the "sea-star" arms of starfish and brittle stars, as well as the rounded shape of Echinozoa, both evolved independently from an ancestor of unknown morphology, but that each "armed" and "rounded" lineages are strictly monophyletic. Too little is known of the basal eleutherozoans and echinoderms to be able to firmly decide for or against any of these hypotheses at present.[1]

The Asterozoa would have to be ranked as a superclass or treated as an unranked clade between the Cryptosyringida and the Eleutherozoa, depending on whether the "Asterozoa" or "Cryptosyringida" hypothesis turns eventually out to be correct.

Emerging research favours the following classification:[2][3]



Crinoids


ELEUTHEROZOA
Asterozoa

Asteroidea



Somasteroidea





Ophiuroids


Echinozoa

Holothuroideas



Echinoids





[edit] Footnotes

  1. ^ Wray (1999)
  2. ^ Smith, A. (2007). "Echinoderms: Attachment, torsion and the origins of a radical new body plan". in Budd, G.E.; Streng, M.; Daley, A.C.; Willman, S.. Palaeontological Association Annual Meeting. Programme with Abstracts. 51. Uppsala, Sweden. http://downloads.palass.org/annual_meeting/2007/palass2007_programme_abstracts.pdf. 
  3. ^ Smith, A.B. (2005). "The pre-radial history of echinoderms". Geological Journal 40: 255–280. doi:10.1002/gj.1018. 

[edit] References





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