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Aetosaurs
Fossil range: Late Triassic
Desmatosuchus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Subclass: Diapsida
Infraclass: Archosauromorpha
(unranked): Crurotarsi
Order: Aetosauria
Lydekker, 1889
Aetosaur Desmatosuchus from Petrified Forest National Park, Arizona (artist's reconstruction). This is likely the animal that made the nests discovered there in 1996.

The aetosaurs (literally "Eagle Lizards") are an extinct clade of heavily armoured, medium- to large-sized Late Triassic herbivorous archosaurs. Two distinct subdivisions of aeotosaurs are currently recognized, the Desmatosuchinae and the Aetosaurinae, based primarily on differences in the morphology of the bony scutes of the two groups (Parker et al., 2008; p. 692).

The head is small relative to the large body, and quite distinctive in shape, being flat and blunt at the front, like the snout of a pig. The chisel-shaped teeth are small and leaf-like, indicating a probable herbivorous diet (Carroll, 1988; p.273), although peg-like teeth and a keratinous snout have been described in at least one species as possible adaptations for feeding on colonial insects (Small, 2002). Study of the braincase indicates that aetosaurs are actually closely related to crocodylomorphs (Gower & Walker, 2002).

As with the Rauisuchia, they had a "pillar-erect" erect limb posture (Heckert & Lucas, 2002). The feet, however, resemble those of the phytosaurs in the retention of primitive characteristics (Carroll 1988 p.273). In other respects, they have a typically crurotarsan (rauisuchian or crocodylian) body and large powerful tail. Although the fore-limbs are much smaller than the hind limbs, all aetosaurs were quadrupeds.

These animals were very heavily armored (most certainly as a defense against predators), with large quadrangular, interlocking bony plates, or osteoderms, protecting the back and sides, belly, and tail (Carroll, 1988 p.273). In life, these plates were probably covered in horn (Colbert, 1969 p.159).

Primitive genera, like the widespread Norian genus Aetosaurus and the Carnian Coahomasuchus (Heckert & Lucas, 1999), tended to be small, about a meter in length. However more advanced forms were larger - about 3 meters in length - with taxa some, such as Typothorax and Paratypothorax, possessing broad turtle-like bodies, and others, like Desmatosuchus, a narrow-bodied genus up to 5 meters long, equipped with large spines over the shoulders, which added to the animal's defensive armament.

At least some aetosaurs built nests and protected their eggs. In 1996, geologist Stephen Hasiotis discovered 220 million-year-old, fossilized, bowl-like nests in Arizona's Petrified Forest, in part of the Chinle Formation. The oldest such nests that have been found belonged to phytosaurs and aetosaurs. The nests are compacted and appear very similar to the nests of the modern day crocodiles who guard their nests. The nests were holes dug in the sand in the bank of an ancient river.[1]

Aetosaur fossil remains are known from Scotland, Germany, Greenland, the southwest and the eastern United States, Argentina, and Madagascar. Since their armoured plates are often preserved, and as they often have a wide geographic distribution, but a relatively short stratigraphic range, aetosaurs can serve, like phytosaurs, as important Late Triassic tetrapod index fossils (Heckert & Lucas, 2002; Lucas, 1998).


Contents

[edit] Classification

[edit] Taxonomy

[edit] Genera

Genus Status Age Location Description Images
Valid.

Late Triassic.

An herbivore that was protected by spikes and armor plating.
Nomen dubium.

Late Triassic.

An aetosaur whose name means "sturdy lizard." It may be a junior synonym of Stagonolepis.

Junior synonym.

  1. N/A.
N/A. Junior synonym of Stagonolepis.
Valid.

Late Triassic.

A small (1.5 m, 5 ft long) but robustly armored herbivore with an upturned snout. Several species have been described. It had a wide range, skeletal remains have been discovered in both Europe and North America. Fossiliized scutes have been discovered in Madagascar as well.
Junior synonym.
  1. N/A.
N/A. Junior synonym of Stagonolepis
Junior synonym. N/A N/A Junior synonym of Stagonolepis.
Valid.
Valid.
Lapsus calami. lapsus calami of Desmatosuchus
Valid. Late Triassic. North America.
Junior synonym. Possible junior synonym of Rioarribasuchus
Valid. Late Triassic. North America.
Valid.
Valid.
Valid.
Valid.

Might not be an aetosaur.

Valid. — possible junior synonym of Heliocanthus
Valid. Late Triassic. North America.
Valid. Late Triassic. Europe, Geopark of Paleorrota Brazil.
Valid. Late Triassic. North America.
Valid.
Valid. Late Triassic. North America.

[edit] Phylogeny

Cladogram from Parker et al. (2008):

Aetosauria 
 Stagonolepididae 
 Aetosaurinae 


Coahomasuchus



Neoaetosauroides




Stagonolepis



Calyptosuchus



Aetosaurus



Aetosauroides




 Typothoracisinae 
 Paratypothoracisini 

Tecovasuchus




Rioarribasuchus



Paratypothorax






Typothorax



"Redondasuchus"





 Desmatosuchinae 


Longosuchus



Lucasuchus



Acaenasuchus



Sierritasuchus




Desmatosuchus





[edit] References

  • Benton, M. J. (2000). Vertebrate Paleontology, 2nd ed. Blackwell Science Ltd
  • Carroll, R. L. (1988). Vertebrate Paleontology and Evolution, WH Freeman & Co.
  • Colbert, E H. (1969). Evolution of the Vertebrates, John Wiley & Sons Inc (2nd ed.)
  • Gower, D. J. and Walker, A. D. (2002). New data on the braincase of the aetosaurian archosaur (Reptilia: Diapsida) Stagonolepis robertsoni Agassiz, Zoological Journal of the Linnean Society 136: 1-7. Archosaurian Anatomy and Palaeontology: Essays in Memory of Alick D. Walker, DB Norman & DJ Gower (eds.)
  • Heckert, A. B and Lucas, S. G, (1999) A new aetosaur (Reptilia: Archosauria) from the Upper Triassic of Texas and the phylogeny of aetosaurs. Journal of Vertebrate Paleontology. 19 (1): 50-68.
  • -- -- (2002), South American occurrences of the Adamanian (Late Triassic: Latest Carnian) index taxon Stagonolepis (Archosauria: Aetosauria) and their biochronological significance, Journal of Paleontology 76 (5): 852-863. online
  • Lucas, S.G. (1998). Global Triassic tetrapod biostratigraphy and biochronology. Paleogeography, Palaeoclimatology, Palaeoecology 143: 347-384.
  • Parker, W.G. et al. 2008. "A new desmatosuchine aetosaur (Archosauria; Suchia) from the Upper Triassic Tecovas Formation (Dockum Group) of Texas." Journal of Vertebrate Paleontology, 28 (2): 692-701.
  • Small, B.J. (2002). Cranial anatomy of Desmatosuchus haplocerus (Reptilia: Archosauria: Stagonolepididae). Zoological Journal of the Linnean Society 136(1): 97-111.

[edit] External links





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